Some stuff you can google on codominance. Roan color.
Roan Cattle
The roan colour can also result in Shorthorn Cattle, but in a case of intermediate inheritance or Codominance. These cattle can not breed true, as this type of intermediate inheritance is also Heterozygous, or hybrid, as two different colour alleles are required form the parents to produce this particular coat coloring.
Breeds of cattle known for roans are the Belgian Blue and Shorthorn. Among the former, coat color may be solid black, solid white, or blue roan; the latter may be solid red, solid white, or red roan. Belgian Blues also typically exhibit spotting patterns,[11] which are genetically separate from roan. As a result, most roan cows exhibit blotches of clearly colored and clearly white hair, with roan patches.[12] Some "cryptic" roan cattle appear solid, but upon close inspection reveal a small roan patch.[13] Roan cattle cannot "breed true" but breeding white cattle to a solid mate will always yield a roan calf. The white color typical of Charolais and White Park breeds is not related to roan.[13]
Roan in Shorthorns and Belgian Blues is controlled by the mast cell growth factor (MGF) gene, also called the steel locus, on bovine chromosome 5.[14] Part of the KIT ligand, this region is involved in many cell differentiation processes. Mast cell growth factor promotes pigment production by pigment cells,[15] and without it, skin and hair cells lack pigment. With two functional MGF genes (homozygous dominant), cattle are fully-pigmented; without any functional MGF genes (homozygous recessive), they are white. MGF-controlled roan occurs when cattle possess one functional and one non-functional MGF gene (heterozygous), resulting in a roughly even mixture of white regions and colored regions.[13]
Codominance
From Wikipedia, the free encyclopedia
Codominance refers to a relationship between two alleles of a gene. It occurs when the contributions of both alleles are visible in the phenotype.
For instance, in the ABO system, the IA and IB alleles are co-dominant in producing the AB blood group phenotype, in which both A- and B-type antigens are made. Another example occurs at the locus for the Beta-globin component of hemoglobin, where the three molecular phenotypes of HbA/HbA, HbA/HbS, and HbS/HbS are all equally detectable by protein electrophoresis. (The medical condition produced by the heterozygous genotype is called an incomplete dominant, see above). For most gene loci at the molecular level, both alleles are expressed co-dominantly, because both are transcribed into RNA.
Co-dominance and incomplete or semi-dominance are not the same thing. For example, in some plant species, pink flowers may be the product of a mixture of red and white pigments (co-dominance on the pigment level, no dominance on the color level), or the result of one allele that produces the usual amount of red pigment and another non-functional allele that produces no pigment, so as to produce a dilute, intermediate pink color (no dominance at either level).